Reproduction and Genetic Selection
Choosing a Bull
Michel A. Wattiaux
The Babcock Institute
The choice of bulls used in the herd today determines what kind of cows will be in production three years in the future. Choosing a bull should be based on the following considerations:
• Prioritize the traits considered for selection and give each trait relative importance-production traits should be three to five times more important than conformation traits
• Use a selection method-the independent culling method or preferably a selection index-to choose a bull based on PTA values
• Do not use reliability to select sires-use reliability to decide how "heavily" to use selected sires
USING PTA FOR SELECTION DECISIONS
The basic purpose of Predicted Transmitting Ability (PTA) is to rank bulls. If a bull has a PTA of +1,000 kg of milk, it does not mean that his daughters will produce 1,000 kg more milk than their herdmates. It does mean, however, that the daughters of this bull have an average milk production 1,000 kg higher than the daughters of the bulls used in the genetic base. Selection should be based on the PTA value of a bull. A common mistake is to use reliability as a criteria for selection. The proper way to use reliability values will be explained later. In terms of selecting sires, two methods are available: the independent culling level and the selection index. Although both methods have advantages and disadvantages, the index method is preferred by geneticists because it usually offers the possibility for greater genetic progress.
Independent culling level
Independent culling level is a method by which the producer sets a minimum value for each trait in the selection program (In the case of calving ease, the criteria is not a minimum, but a maximum value for percentage of difficult calving). Sires above the minimum for all traits are considered for selection. For example, assuming that two traits of importance are PTA for milk yield and PTA for protein yield, one might decide to choose from among sires with a PTA milk yield greater than 2,250 pounds and a PTA protein yield greater than 65 pounds. As illustrated in Figure 1 only two bulls in the US population in January 1995 would meet these standards.
Advantages
This is the simplest method to identify bulls that meet the goals of a selection program.
Disadvantages
The first difficulty in using the independent culling level is to set the (minimum) standards. A sire may be rejected for failure to meet a standard, even if it is only by a few pounds, while all other traits may far exceed the minimum standards (Figure 1). The need to update the standards periodically is another drawback of independent culling levels. When sires are selected with this method, genetic progress and the change in genetic base may affect which sire becomes the most desirable. If a set of standards has been used for several years, more bulls tend to be in the acceptable group because of genetic progress. Other criteria must be found to select the desired sire or the standards have to be adjusted periodically. In addition, when the genetic base is changed, only very few (if any) of the bulls might meet the standards; again, the standards need to be reviewed and adjusted accordingly.
Figure 1: Selection of sires using the independent culling level
Figure 1: Selection of sires using the independent culling level
Selection index
Use of a selection index allows the ranking of bulls based on a value calculated by giving each selected trait a "relative weight." The "weight" represents the importance that a producer chooses to give to a particular trait. The actual price for milk components, for example, may be used as a weighing factor. In this case, the index would have a currency unit (dollars, francs, etc.). However, the absolute value of an index is really of little importance. Bulls should be ranked according to the most appropriate index, and the highest bulls on the index should be used regardless of the value of the index itself. In other words, there is no significance to setting minimum standards for the value of an index.
Advantages
An index allows identification of bulls that best fit the overall genetic goal without focusing on any trait in particular. Indices force producers to consciously evaluate the traits they want to emphasize and formulate a specific plan for maximizing them in the herd. Once an index is built properly, selection of sires is much simpler than independent culling levels because the best bulls, simply, are the ones that rank at the top of the list. In addition, an index is an objective method to give credit to sires that may be rejected using an independent culling method because they fall short of the standard for one trait.
Disadvantages
Indices are difficult to build because it is difficult to choose the traits to include and the weight to assign to each trait. Many "pre-calculated" indices are now available. Which of the available indices reflects the priority set for a herd is a difficult question to answer. Some indices give an important weight to conformation traits. In other indices, the economic weight given to yield traits is based on the structure of current milk prices in the United States. The interpretation of these indices is difficult for dairy farmers of other countries because, clearly, the pricing of the milk varies widely from one country to another.
How to build a selection index that reflects expected increases in gross income
As an example, let us assume that we are building an index for a market that has the following milk pricing structure. Let us assume that the milk price is 12.2 (any currency unit) per kg for milk containing 3.5% fat and 3.2% protein. The processing plant pays 0.150 currency units per 0.1% of fat (or per gram of fat) and 0.300 currency units per 0.1% of protein (or per gram of protein). The relative weight given to milk yield, protein yield and fat yield may be calculated as follows:
• Value of 1 gram of fat = 0.150 currency units, value of 1 kg of fat = 150 currency units, value of the 35 g of fat in 1 kg of milk = 0.150 x 35 = 5.25 currency units
• Value of 1 gram of protein = 0.300 currency units, value of 1 kg of protein = 300 currency units, value of the 32 g of protein in 1 kg of milk = 0.300 x 32 = 9.60 currency units
• Value of 1 kg of milk without fat and protein = 12.2 - 5.25 - 9.60 = -2.65 currency units
Thus we can calculate an index that will reflect the expected increase in gross income of the daughters of a sire based on his PTA for milk, protein and fat yields. Let us call this index PTA-AGI (Predicted Transmitting Ability of Added Gross Income). We use the terminology gross income because, as cows produce more milk, they eat more; however, the extra cost of feeding is not included in our index.
PTA-AGI = (-2.65 x PTA kg of milk) + (300 x PTA kg of protein) + (150 x PTA kg of fat).
For example a sire with milk PTA of 800 kg, a protein PTA of 20 kg, and a fat PTA of 38 kg, would have a PTA-AGI index of:
PTA-AGI = (-2.65 x 800) + (300 x 20) + (150 x 38) = 9,580 currency units
HOW MANY BULLS SHOULD BE USED IN A HERD?
The number of bulls and the strategy of selecting bulls may differ as a function of:
• Herd size
• Bull's reliability
• The inclination of the farmer to take measured risks. free of serious mastitis
When young sires are selected, the purchase of semen should be limited to only a few units per bull so as to spread the risk. As reliability increases, semen purchases per bull can increase. There is little reason to breed more than 15 to 20% of a herd to any one sire, even if the sire has a PTA with a high reliability. In other words the minimum would be to select at least three sires for every 50 cows in the herd. Diversification is a safeguard against any unforeseen problems that may arise as the result of using any one bull heavily in a herd.
USE RELIABILITY TO DECIDE HOW MUCH TO INVEST IN A SIRE
Reliability indicates the accuracy of genetic evaluations. Often producers use reliability as a criteria for selection. Actually, reliability should never be used in selecting sires; however, once sires have been selected, reliability should be used to decide the intensity of usage (i.e., quantity of semen to purchase). The number of available sires is large and it is tempting to reject those for which reliability of PTA is low. However, using reliability as a selection criteria is likely to limit genetic gain.
Figure 2: Example of change in PTA of 20 sires as reliability increases from 70% to 99%; each sire is identified by a rectangle labeled from 1 to 20 and the distribution of their daughters is illustrated by the horizontal bell-shaped curves.
Figure 2: Example of change in PTA of 20 sires as reliability increases from 70% to 99%; each sire is identified by a rectangle labeled from 1 to 20 and the distribution of their daughters is illustrated by the horizontal bell-shaped curves.
The true genetic value of sires that have a PTA with high reliability is very certain and is not likely to change over time. For some producers, this feature may be important because it guarantees that the genetic merit of the daughters is predicted to be in a narrow range around the PTA. However, high reliability might be considered by others as lack of opportunity. Younger bulls usually have lower reliability because of the limited numbers of daughters in their progeny test. Yet, bulls with the highest genetic merit are among the bulls being progeny tested; we just do not know which ones they are. So how can one take full advantage of the genetic superiority of certain (young) bulls, despite the uncertainty of their PTA?
We know that a PTA accompanied by a low reliability is likely to change, but we cannot predict ahead of time whether the PTA will go up or down. Indeed, when two sires have the same PTA, it is actually more risky to buy a lot of semen from a sire with lower reliability.
If instead of focusing on one young sire, we focus on a group of young sires, the risks will change drastically. Let us consider the future of a group of young sires with high PTAs and low reliability (Figure 2). In general, any time the PTA of a sire decreases, the PTA of another increases. In a group, the bulls that decline in PTA usually are counterbalanced by bulls that increase, for an average of near zero. As a result, the overall PTA average of a group of young sires will remain unchanged over time because the decline in PTA of some of them will be compensated by the increase in the PTA of others (Figure 2). In other words, the best strategy to minimize the risk of low reliability and still maximize potential genetic gain is to focus on low reliability sires as a group. The strategy should be to buy some semen from many young sires as opposed to buying a lot of semen from only a few young sires. By purchasing a few units of semen from several different, low reliability bulls, the chances of milking a large number of daughters from any one bull that may decline in proof is limited; and the chances of milking a few daughters from any one bull that may increase greatly in proof is maximized.
Not all bulls/bucks from the same herd will have the same rate for production.Some will be higher and some lower.The reliability factor is really hard to figure out for bucks over cattle because of the fact.In the west only a handful of bulls are responsible for the majority of calves born,realiability is easier to measure.Buck goats are so spread out that tracking any single buck is difficult.Any buck with 30 daughters in 30 different herds in the Philippines is still some time away.The importance of using more than 1 breeding buck in your herd and by observations and data collection can help the breeder/producer keep track for reliability.Not all bucks born on the farm will score high for reliability which is what that buck transfers in traits needed,two out of three times to his offsprings.
This gets into some really advanced breeding.